Gating charge displacement in a monomeric voltage-gated proton (H v 1) channel

Abstract

H v 1 proton channels, since their open probability increases with depolarization and low pH, are fundamental in sustaining the suitable pH gradient for cell survival. Here, we have characterized the gating current elicited by the monomeric mutant proton channel N264R with the aim of understanding the voltage-dependent processes that control channel opening. Gating currents precede ion currents, indicating that a large fraction of gating charge is displaced before H v 1 opening. The voltage sensor displacements are complex and consist of numerous well-defined states. However, most of the charge is displaced in a single transition that probably leads to channel opening. The positively charged arginine in the N264R channel promotes gating charge trapping in addition to blocking the proton currents.

Document Details

Document Type
Pub Defense Publication
Publication Date
Aug 20, 2018
Source ID
10.1073/pnas.1809705115

Entities

People

  • Alan Neely
  • Carlos E. Gonzalez
  • Emerson M. Carmona
  • H Peter Larsson
  • Osvaldo Alvarez
  • Ramon Latorre

Organizations

  • Air Force Office of Scientific Research
  • CONICYT
  • National Fund for Scientific and Technological Development
  • National Institute of General Medical Sciences
  • University of Miami
  • University of Valparaíso

Tags

Readers

  • Molecular and Cellular Biochemistry
  • Pulsed Power and Plasma Physics.
  • Radio communications and signal processing.