Structural basis for polarized elongation of actin filaments

Abstract

Eukaryotic cells utilize actin filaments to move, change shape, divide, and transport cargo. Decades of experiments have established that actin filaments elongate and shorten significantly faster from one end than the other, but the underlying mechanism for this asymmetry has not been explained. We used molecular dynamics simulations to investigate the structures of the actin filament ends in the ATP, ADP plus γ-phosphate, and ADP nucleotide states. We characterize the structures of actin subunits at both ends of the filament, explain the mechanisms leading to these differences, and connect the divergent structural properties of the two ends to their distinct polymerization rate constants.

Document Details

Document Type
Pub Defense Publication
Publication Date
Nov 16, 2020
Source ID
10.1073/pnas.2011128117

Entities

People

  • Gregory A. Voth
  • Harshwardhan Katkar
  • Steven Z Chou
  • Thomas D. Pollard
  • Vilmos Zsolnay

Organizations

  • National Institute of General Medical Sciences
  • United States Department of Defense
  • University of Chicago
  • Yale University

Tags

Readers

  • Economics
  • Electrical Engineering
  • Molecular and Cellular Biochemistry