Pupil Size Encodes Uncertainty during Exploration

Abstract

Exploration is an important part of decision making and is crucial to maximizing long-term rewards. Past work has shown that people use different forms of uncertainty to guide exploration. In this study, we investigate the role of the pupil-linked arousal system in uncertainty-guided exploration. We measured participants' (n = 48) pupil dilation while they performed a two-armed bandit task. Consistent with previous work, we found that people adopted a hybrid of directed, random, and undirected exploration, which are sensitive to relative uncertainty, total uncertainty, and value difference between options, respectively. We also found a positive correlation between pupil size and total uncertainty. Furthermore, augmenting the choice model with subject-specific total uncertainty estimates decoded from the pupil size improved predictions of held-out choices, suggesting that people used the uncertainty estimate encoded in pupil size to decide which option to explore. Together, the data shed light on the computations underlying uncertainty-driven exploration. Under the assumption that pupil size reflects locus coeruleus-norepinephrine neuromodulatory activity, these results also extend the theory of the locus coeruleus-norepinephrine function in exploration, highlighting its selective role in driving uncertainty-guided random exploration.

Document Details

Document Type
Pub Defense Publication
Publication Date
Jan 01, 2023
Source ID
10.1162/jocn_a_02025

Entities

People

  • Deshawn Chatman Sambrano
  • Elizabeth A. Phelps
  • Emily Dial
  • Haoxue Fan
  • Samuel J Gershman
  • Taylor Burke

Organizations

  • Air Force Office of Scientific Research
  • Harvard University
  • Massachusetts Institute of Technology
  • National Institute on Drug Abuse

Tags

Readers

  • Computational Modeling and Simulation
  • Neural Network Machine Learning.
  • Vision Science/Vision Psychology/Cognitive Neuroscience.