Membrane Voltage Effects on Proton Transport by a Yeast H+ -ATPase

Abstract

The objective of this two year project was to identify protein structure domains participating in proton transport and membrane voltage interactions by the plasma membrane H(+)-ATPase from Saccharomyces cerevisiae. H+-ATPase mutants (pmal) were generated by random and site-directed mutagensis techniques that cause a deploarization of the cellular membrane potential. All pmal mutant enzymes were active in proton transport although one mutant Gly158-- >Asp, appeared to be partially uncoupled from ATP hydrolysis. Three loci, one within a putative transmembrane domain (GLY158) and the other two (Ser368, Pro640) within putative membrane/cytoplasmic interface domains, were found to cause the most prominent effect on cellular membrane potential. To better understand how membrane voltage effects the H(+)-ATPase, a new procedure was developed to produce large and sustained membrane potentials in reconstituted proteoliposomes. The results of this project will serve as a foundation for probing electrogenic proton transport by the H(+)-ATPase with the eventual goal of developing a structural model for ion translocation. Keywords: Cations, Phosphorus Hydrolases.

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Document Details

Document Type
Technical Report
Publication Date
Jul 31, 1989
Accession Number
ADA211990

Entities

People

  • David S. Perlin

Organizations

  • Public Health Research Institute

Tags

Communities of Interest

  • Biomedical

DTIC Thesaurus Topics

  • Amino Acids
  • Cell Membrane
  • Cells
  • Chemistry
  • Enzymes
  • Hydrolysis
  • Measurement
  • Membrane Potentials
  • Membranes
  • Military Research
  • Osmotic Pressure
  • Public Health
  • Sensitivity
  • Steady State
  • Transport Properties
  • Transport Ships
  • Vanadium Compounds

Fields of Study

  • Biology

Readers

  • Molecular and Cellular Biochemistry